The film continues immediately after the events of Casino Royale with Bond driving from Lake Garda to Siena, Italy. With the captured Mr. White in the luggage compartment of his car, Bond is attacked by chasing henchmen. After evading his pursuers, Bond and M interrogate White regarding his secretive organisation, Quantum. It becomes apparent that MI6 knows little to nothing about Quantum. Bond pockets a picture of his former lover Vesper Lynd and her boyfriend, Yusef Kabira, after M tells Bond that Kabira is still alive. M's bodyguard, Mitchell, is revealed as a traitor, attacking M and allowing White to escape; Bond chases Mitchell across Siena and kills him. Following a forensic investigation into Mitchell's apartment, Bond heads to Haiti to track down and kill Mitchell's contact. Bond learns that henchman Slate was sent to kill Camille Montes at the behest of her lover, Dominic Greene, the chairman of an ecological organization called Greene Planet. While observing her meeting with Greene, Bond learns that Greene is helping the Bolivian general Medrano (who murdered Camille's family) overthrow his government in exchange for a seemingly barren piece of desert.Greene has Camille escorted away on Medrano's boat to "sweeten" their deal, but Bond rescues her. Bond then follows Greene to a private jet, which flies him to a performance of Tosca at Lake Constance, Austria. Bond infiltrates Quantum's meeting at the opera, and a gunfight ensues in a restaurant. A bodyguard of Guy Haines, an advisor to the British Prime Minister, is accidentally killed, and M has Bond's passports and credit cards revoked because she believes Bond has killed too many potential sources of information. Bond travels to Italy to reunite with his old ally René Mathis, whom he convinces to accompany him to La Paz. On the flight over, Bond drinks his signature drink and broods over Vesper; he feels betrayed and heartbroken. They are greeted by Strawberry Fields, an MI6 field operative from the British Consulate, who demands that Bond return to the UK on the next available flight. Bond disobeys and seduces her in their hotel suite.
Bond meets Camille again at a ecological fund-raiser being held by Greene, where she is busy spoiling Greene's party by pointing out his hypocrisy and lies to wealthy donors. Bond and Camille leave hastily together, but are pulled over by the Bolivian police. The police order Bond to open the luggage compartment of his vehicle, revealing a bloodied Mathis. As Bond lifts Mathis out of the vehicle, the policemen open fire and fatally wound Mathis. After Bond subdues the police, he has a moment of relative tenderness with Mathis as the dying man asks Bond to stay with him. Mathis tells Bond that Vesper "gave everything" for him; his dying wish is that Bond forgive Vesper, and that he forgive himself. After depositing Mathis' body in a waste container, Bond and Camille drive to Greene's intended land acquisition in the Bolivian desert and survey the area in a Douglas DC-3 plane. They are intercepted and shot down by an Aermacchi SF.260 fighter and a Bell UH-1 Iroquois helicopter. They escape from the crippled plane by parachuting into a sinkhole. While escaping the cave, Bond and Camille discover Quantum is blockading Bolivia's supply of fresh water, normally flowing in subterranean rivers, by damming it. Bond also discovers that Camille has spent years plotting revenge on General Medrano for the murder of her family. The duo return to La Paz, where Bond meets M and learns Quantum murdered Fields by drowning her in petroleum. Believing that Bond has become a threat to both friend and foe, and acting under higher orders, M orders him to disarm and end his activities in Bolivia. Bond escapes, defying M's orders to come in. M tells her men to watch him because she thinks he is "on to something." Even though he has gone rogue, M confesses that she still has faith in him, and that he is "still [her] agent."
By this point, both the American and British governments have agreed to work with Greene, because they think he has control of vast supplies of oil in Bolivia. Bond meets American CIA agent Felix Leiter (a colleague from the high-stakes poker game in Casino Royale ) at a local bar, who, like Bond, thinks his government is on the wrong track. Leiter discloses that Greene and Medrano will meet at an eco-hotel in the Bolivian desert.
At the meeting in the eco-hotel, Greene threatens Gen. Medrano into signing a contract granting Greene's company an overpriced proprietary utilities contract in Bolivia. After the meeting, Bond attacks and kills the Colonel of Police for betraying Mathis, and sets off a chain of explosions in the hotel when a hydrogen fuel tank is hit by an out of control vehicle. Camille busts up Medrano's attempted rape of his servant girl, and after a fight (during which Medrano tells Camille that she has the same frightened look her mother had before he killed her), Camille kills Medrano. Bond rescues Camille from the burning building, and captures Greene. After interrogating him, he leaves Greene stranded in the middle of the desert with only a can of motor oil. Bond tells him that he bets Greene will make it 20 miles across the desert before he considers drinking the oil, contrasting the resources of oil and water. Bond drives Camille to a train station, where she muses on what life holds for her now that her revenge is complete. They kiss briefly but passionately before she departs.
Bond goes to Kazan, Russia, where he confronts Vesper Lynd's former lover Yusef Kabira. Yusef is a member of Quantum who seduces high-ranking women with valuable connections, getting them to give up government assets as ransom for himself in fake kidnappings where he is supposedly held hostage. He is attempting to do the same with Canadian agent Corinne Venneau, even giving her the same kind of necklace he gave Vesper. Surprising them at Yusef's apartment, Bond tells Corinne about Vesper and advises her to alert the Canadian Security Intelligence Service. Bond leaves Yusef's apartment and is confronted by M who is surprised that Bond did not kill Yusef, but rather left him alive for questioning. M reveals that Leiter has been promoted at the CIA to replace Beam, and that Greene was found in the desert, shot dead and with motor oil in his stomach. Bond doesn't volunteer any information on Greene, but tells M that she was right about Vesper. M then tells Bond that MI6 needs him and fully reinstates him as an agent. Bond walks off into the night telling M that he never left. As he leaves, he drops Vesper's necklace in the snow.
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