Manny the woolly mammoth, Sid the sloth, Diego the saber-toothed tiger, and the hapless prehistoric squirrel/rat known as Scrat are still together and enjoying the perks of their now melting world. Manny may be ready to start a family, but nobody has seen another mammoth for a long time; Manny thinks he may be the last one. That is, until he miraculously finds Ellie, the only female mammoth left in the world. Their only problems: They can't stand each other--and Ellie somehow thinks she's a possum! Ellie comes with some excess baggage in the form of her two possum "brothers"-- Crash and Eddie, a couple of daredevil pranksters and cocky, loud-mouthed troublemakers. Manny, Sid and Diego quickly learn that the warming climate has one major drawback: A huge glacial dam holding off oceans of water is about to break, threatening the entire valley. The only chance of survival lies at the other end of the valley. So our three heroes, along with Ellie, Crash and Eddie, form the most unlikely family--in any "Age"-- as they embark on a mission across an ever-changing, increasingly dangerous landscape towards their salvation. While Skrat the Saber Toothed Squirrel is still ineffectually trying to regain his precious acorn, the misfit trio of Manny the Mammoth, Sid the Sloth and Diego the Sabretooth Tiger have settled down in an isolated valley with numerous other animals. However, the trio discover that the ice wall surrounding the valley is barely holding back a massive body of water behind it and it's melting state threatens to break and flood the valley. With their one chance of survival being a boat at the other end of the valley, the trio follow the desperate exodus there. Along the way, they meet Ellie, a female mammoth who is convinced she's a opossum like her brothers. While the strange group continues the trek, they must learn to get along even as Manny struggles to find some connection to this strange female who may be the only other one of his kind.
When the valley where they live is threatened by the melt of the surrounding ice, the mammoth Manny, the sloth Sid and the saber-toothed tiger Diego advise the other animals to look for shelter outside the place. While escaping, the trio meets Ellie, a female mammoth that believes she is an opossum like her two mischievous brothers. Meanwhile, the squirrel Scrat tries to retrieve his walnut in the most unusual situations.
Manny, Sid, and Diego are currently living in a large valley surrounded by an enormously high ice wall on all sides. When a water park begins to form, the trio discovers that the ice wall is actually a wall that is barely holding a massive body of water that could flood the valley to nearly a mile underwater. A vulture tells them that there is a boat at the other end of the valley that may save them all, but they only have 3 days to make it or die. Manny is having trouble facing the fact that he may be the last mammoth left. Along the way, they meet Ellie, a mammoth who thinks she is a possum, and her possum brothers Crash and Eddie. As they begin to travel together, Manny learns, with help from Diego and Sid, that having this new mammoth with them, may not be as bad as Manny makes it out to be while they try to escape the oncoming and continually threatening flood. During their journey, Scrat the sabre-toothed squirrel has his own hilarious adventures that eventually leads to something else entirely.
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